Category Archives: Nursery News

Nursery Knowledge: Plant Hydraulic Physiology

Lloyd Nackley

Unlocking Nature’s Water Secrets for Greener Futures, Part 1

TL;DR Plant hydraulics unravels the journey of water within plants, aiding tree health, nursery production, urban forest management, and climate resilience. 🌿🌿

Last month, we delved into the fascinating world of soil hydraulics, exploring how water moves beneath our feet. In this post, we’re staying within the realm of water movement but shifting our focus to a different dimension of nature – plants. Prepare to journey through the intricate pathways of plant hydraulic physiology, where we uncover the secrets of how trees and other woody plants manage water, adapt to challenging conditions, and ultimately contribute to a greener, more sustainable
world.


Plant hydraulic physiology is all about how water moves through plants. Scientists study this to understand how trees and other woody plants react when they have enough water or not enough. This knowledge helps us figure out how different ways of growing plants in nurseries affects their growth. People have known for a while that this field is important for plants in forests. But now, thanks to recent discoveries by this lab and others, this amazing field of science is being applied to
nurseries and other horticultural production systems. In this summary, I will explain the basic ideas about how water moves through
plants , how it connects to their structure and how they work. With this knowledge, scientists, nursery workers, and people who care for forests can ensure they grow strong, healthy trees that can handle harsh conditions when planted outside.

UNDERSTANDING WATER MOVEMENT IN PLANTS
Let’s start by talking about how water moves in plants. Imagine it’s like water moving through a hose in your garden. We can measure this flow of water using something called “flow rate,” which is just how much water moves in a certain amount of time.

We use units like gallons per minute or liters per minute to measure it. For example, think about a water hose in your garden. If you want to know how much water it sprays out in a minute, that’s its flow rate. Now, here’s something interesting: the size of the pipe or hose matters. A big hose can let a lot more water flow through than a tiny one. In fact if you double the diameter of a hose it can allow 4 times the flow of the smaller diameter hose. Plants have tiny water pipes called “xylem.”

XYLEM CONDUCTANCE
Okay, now let’s talk about “conductance.” Think of it as how easy or hard it is for water to move through something. For plants, this refers to how easily water can travel through their pipes. We usually keep the pressure the same, like when you use a hose with a constant water pressure. This helps make sure the plants get water evenly. Lastly, there’s something called “conductivity.” It’s like a fancy version of conductance but scaled to the size of different parts of the plant. It helps us compare how different parts of the plant move water. For example, we might want to know how water moves through the stem compared to the roots.

Now, here’s where it gets cool: in plants, water doesn’t get pushed like in your garden hose. It gets pulled up by something called “tension.” This happens because plants lose water from their leaves when it evaporates. Imagine a plant sipping water through a straw from the soil. When the water evaporates from the leaves, it creates tension, like a vacuum pulling water up the plant. This is how water can move up the tallest trees. So, we measure something called “water potential” to understand this tension. It tells us how much “pull” the plant has on the water. When there’s a difference in water potential between different parts of the plant, it’s like a driving force that makes water move from where there’s less pull to where there’s more. This helps water move up from the roots to the leaves, even against gravity. We call this whole process the “Soil-Plant-Atmosphere Continuum,” but you can just think of it as how plants drink water.

And that’s the basics of how water moves in plants!

Nursery Knowledge: What’s in the Pot?

Exploring Stratified Substrates and Soil Hydraulics in Agricultural Science

Lloyd Nackley

Nursery science researchers have embarked on a journey to harness the principles of soil hydraulics to reshape container production practices. In the past couple of years, Dr. Chris Criscione and Dr. Jeb Fields and others released a series of articles that showed that stratifying pine bark can serve as a substitute for peat-based media in floriculture and bark-based woody plant production. Through layering premium floriculture media over cost-effective pine bark within containers, they reduced reliance on peat. In their study focused on Petunia hybrid ‘Supertunia Honey’, the stratified substrates yielded crops of comparable size and quality, along with enhanced root productivity. Their work also showed superior performance of bark:coir substrates in a stratified setup, even when subjected to reduced irrigation when producing Loropetalum chinense ‘Ruby’ liners. Positive microbial communities in stratified systems further aided in mitigating water stress. Furthermore, Red Drift® rose plants grown in stratified substrates exhibited equal or superior crop growth despite receiving less controlled-release fertilizer. This suggests potential for reducing fertilizer and irrigation rates while upholding crop
quality, offering a sustainable avenue for containerized crops. This blog post aims to shed light on stratified substrates and also provide insights into ongoing projects at the Nackley Lab that are delving into this innovative frontier of nursery science.

image shows containers of growing medium of different textures
Image: mixing coarse and fine substrates in containers to address soil stratification

Soil hydraulics delves into the intricate world of water movement within soils—how water traverses the soil matrix, its distribution, and its intricate interactions with soil particles and structure. It’s a field dedicated to understanding the physics governing water’s journey through soils, and this knowledge carries significance for agriculture, environmental science, hydrology, and civil engineering. Soil hydraulics guides irrigation strategies, shapes drainage designs, and fuels sustainable land management practices.

image shows saturated and unsaturated soils in comparison
Image: water in saturated and unsaturated soils

Central to soil hydraulics is the concept of matric potential, a critical factor in irrigation management and plant-available water. Matric potential captures the degree to which soil particles retain water due to molecular attractions—it’s essentially the “stickiness” of water to soil particles. When soil isn’t completely saturated, minuscule air-filled spaces exist between particles. Water molecules adhere to these particles, creating capillary forces that coax water upwards. The strength of the bond between water and soil particles determines the matric potential, effectively influencing water availability to plants.

Image shows layering of the 3 soil phases: gaseous, liquid and solid
Image: the gaseous, liquid and solid phases of soils

Two other essential players in the realm of soil hydraulics are soil texture and structure. Envision soil as an intricate mosaic, with particles coming together to form aggregates, shaping distinct pathways and chambers. This arrangement, referred to as soil structure, creates macropores—
akin to express highways—through which water can flow rapidly. At the same time, micropores, reminiscent of narrow alleyways, gently retain water against gravity’s pull, acting as reservoirs
for plant roots during dry spells. Additionally, soil structure influences permeability, determining how efficiently water infiltrates and the risk of surface runoff.

Soil texture, on the other hand, hinges on the proportions of sand, silt, and clay particles in the soil. Each particle type comes with distinct traits that shape water dynamics. Coarser-textured soils with more sand have ample space between particles, allowing water to move quickly with
less retention. Finer-textured soils, rich in silt and clay, boast smaller gaps between particles, leading to slower water movement and higher water-holding capacity.

image shows the soil texture triangle
Image: The Soil Texture Triangle

However, when soil is confined to pots or containers, it undergoes transformations in its natural structure due to various container-related factors. The restricted space of a pot, contrasting with the expanse of natural soil, can lead to the erosion of soil structure. Aggregates—clusters of particles forming pores and pathways—can deteriorate over time due to limited expansion room. This confined space often results in compaction, where soil particles compress closely, reducing essential air-filled pores needed for root growth and water movement. The watering practices specific to potted plants can contribute further by compacting soil particles as water occupies gaps. The absence of natural soil organisms and plant roots within containers hampers the
maintenance of soil structure. Repeated disturbances, like transplanting or repotting, can exacerbate these structural changes. To counter these effects, selecting appropriate potting mixes that retain structure, incorporating organic matter for improved aeration and water retention, and being mindful of compaction during planting and watering are recommended.

image shows a hydrangea in a pot with drip irrigation
Image: Hydrangea in pot with drip emitter irrigation

Traditionally, the nursery field has focused on creating homogeneous potting mixes that maintain structure while offering suitable hydraulic properties. Classic blends often comprise bark, coir, peat, perlite, vermiculite, and pumice. However, a recent shift in focus has led scientists to explore how layering media can simulate natural soil hydraulics—an approach known as stratified substrates.

coarse substrate
Coarse substrate
fine substrate
Fine substrate

Stratified substrates involve arranging potting media of varying textures in layers within a single container. This structured layering entails placing coarser-textured substrates at the bottom and finer-textured ones on top, mimicking natural soil layers. This technique aims to influence water
movement, nutrient distribution, and hydraulic behavior within the confined environment of a container. By borrowing from the stratification seen in the ground, stratified substrates strive to
optimize resource efficiency, plant growth, and root development in controlled settings like potted plants.

Some may draw parallels between stratified substrates and the practice of placing rocks or gravel at the bottom of larger plant pots. While both concepts involve layering materials, there are distinctions. The practice of adding gravel or rocks to enhance drainage in larger pots shares a
kinship with stratified substrates. However, it doesn’t replicate the comprehensive layering dynamics seen in stratified substrates. Adding gravel mainly addresses drainage concerns without fully incorporating the layered hydraulic principles inherent to stratified substrates.

image shows 2 piles of substrates by texture
Image: fine (l) and coarse (r) substrate, side by side

Dr. Chris Criscione, in collaboration with the Dr. Jeb Fields group at Louisiana State University, has been at the forefront of investigating stratified substrates in containerized plant growth. Their research delves into how layering different potting media textures can enhance water retention, nutrient availability, and overall plant performance. The studies highlight promising outcomes, such as heightened root productivity, improved growth, and enhanced quality under stratified
conditions compared to conventional substrates. This technique holds potential for bolstering sustainable crop cultivation within controlled environments.

gravel pad with pots of hydrangea growing
Image: potted hydrangeas on the gravel pad at NWREC

Nevertheless, it’s crucial to acknowledge that findings from studies conducted in one geographic region—such as the Southeastern USA—may not seamlessly extrapolate to other areas with distinct climates and environmental conditions, like the Pacific Northwest. Climate, temperature, humidity, and other factors can significantly influence plant growth and water dynamics. Given this variation, research in regions like the Pacific Northwest, such as Oregon, is crucial. The
unique environmental factors there, including cooler temperatures and higher rainfall, can impact water movement, nutrient availability, and plant response to stratified substrates. Bark-based substrates, common in some areas, may behave differently in terms of water retention and
drainage in regions with distinct soil compositions. To address this, the Nackley Lab initiated a collaboration with Dr. Fields and others in 2022, planning to explore the impacts and benefits of stratifying substrates in Nursery production. The first stratified substrate experiment was
launched in 2023 at Oregon State University North Willamette Research and Extension Center (NWREC), aiming to reduce resource demand and provide insights into the effectiveness of stratified substrates in that context, contributing to more informed decision-making for nursery production and horticulture practices in the region.

More Information:

Evaluating Stratified Substrates Effect on Containerized Crop Growth under Varied Irrigation Strategies

Root Exploration, Initial Moisture Conditions, and Irrigation Scheduling Influence Hydration of Stratified and Non-Stratified Substrates

Single-screen Bark Particle Separation Can be Used to Engineer Stratified Substrate Systems

Stratified Substrates Can Reduce Peat Use and Improve Root Productivity in Container Crop Production