OHRC Publications

Araki, H., Ardren, W.R., Olsen, E., Cooper, B. and Blouin, M.S., 2007. Reproductive success of captive‐bred steelhead trout in the wild: evaluation of three hatchery programs in the Hood River. Conservation Biology, 21(1), pp.181-190.

Araki, H., Berejikian, B.A., Ford, M.J. and Blouin, M.S., 2008. Fitness of hatchery‐reared salmonids in the wild. Evolutionary Applications, 1(2), pp.342-35

Araki, H., Cooper, B. and Blouin, M.S., 2009. Carry-over effect of captive breeding reduces reproductive fitness of wild-born descendants in the wild. Biology Letters, 5(5), pp.621-624.

Armstrong, M E., Minkoff, D., Dittman, A.H., May, D., Moody, E.K., Quinn, T.P., Atema, J., and W.R Ardren. 2022. Evidence of an olfactory imprinting window in embryonic Atlantic salmon. Ecology of Freshwater Fish, 31, 270–279

Auld H.L., Jacobson, D.P., and M.A. Banks. 2021. Differences in mate pairings of hatchery- and natural-origin coho salmon inferred from offspring genotypes. Integrated Organismal Biology 3(1): obab020.

Auld, H.L. Noakes, D.L.G, and M.A. Banks. 2019. Advancing mate choice studies in salmonids. Reviews in Fish Biology and Fisheries.

Bellinger, M.R., Wei, J., Hartmann, U., Cadiou, H., Winklhofer, M., and M.A. Banks. 2022. Conservation of magnetite biomineralization genes in all domains of life and implications for magnetic sensing. Proceedings of the National Academy of Sciences 119 (3) e2108655119.

Bellinger M.R., Wei, J., Hartmann, U., Cadiou, H., Winklhofer, M., and M.A. Banks. 2022. Reply to Monteil et al.: Universal presence of core magnetite biomineralization genes points to ancient symbiosis. Proceedings of the National Academy of Sciences 119 (39) e2210188119.

Blouin, M.S., Wrey, M.C., Bollmann, S.R., Skaar, J.C., Twibell, R., and C. Fuentes. 2021. Offspring of first-generation hatchery steelhead trout (Oncorhynchus mykiss) grow faster in the hatchery than offspring of wild fish, but survive worse in the wild: Possible mechanisms for inadvertent domestication and fitness loss in hatchery salmon. PLOS One.

Christie, M.R., Ford, M.J. and Blouin, M.S., 2014. On the reproductive success of early‐generation hatchery fish in the wild. Evolutionary Applications, 7(8), pp.883-896.

Christie, M.R., Marine, M.L., French, R.A. and Blouin, M.S., 2012. Genetic adaptation to captivity can occur in a single generation. Proceedings of the National Academy of Sciences, 109(1), pp.238-242.

Church, M.R., Ebersole, J.L., Rensmeyer, K.M., Couture, R.B., Barrows, F.T. and Noakes, D.L., 2009. Mucus: a new tissue fraction for rapid determination of fish diet switching using stable isotope analysis. Canadian Journal of Fisheries and Aquatic Sciences, 66(1), pp.1-5.

Cogliati, K.M., Noakes, D.L., Khan, F., Sharpe, C.S. and C.B. Schreck. 2022. Producing wild fish phenotypes in hatchery-reared fish. Environmental Biology of Fishes, pp.1-22.

Cogliati, K.M., Scanlan, M.M., Self, K.E., Schreck, C.B. and Noakes, D.L.G. 2022. Environmental conditions influence exploration, antipredation behavior, and fin condition in juvenile Chinook salmon (Oncorhynchus tshawytscha). Environmental Biology of Fishes, pp.1-16.

Cogliati, K.M., Unrein, J.R., Sealey, W.M., Barrows, F.T., Hakanson, O., Chitwood, R., Noakes, D.L.G, and C. B. Schreck. 2019. Low-Lipid Diets Fed at Reduced Ration: Effects on Growth, Body Composition, and Survival of Juvenile Chinook Salmon. Journal of Fish & Wildlife Management 10: 500-509.

Cogliati, K.M., Herron, C.L., Noakes, D.L.G., and C.B. Schreck. 2019. Reduced stress response in juvenile Chinook Salmon reared with structure. Aquaculture 504:96-101.

Cogliati, K.M., Unrein, J.R., Stewart, H.A., Schreck, C.B., and D.L.G. Noakes. 2018. Egg size and emergence timing affect morphology and behavior in juvenile Chinook Salmon, Oncorhynchus tshawytscha. Ecology and Evolution 8(1): 778-789.

Cogliati, K.M., Unrein, J.R., Schreck, C.B., and Noakes, D.L.G. 2019. Rearing environment affects spatial learning in juvenile Chinook salmon Oncorhynchus tshawytscha. Journal of Fish Biology 95: 870-880.

Cole, K. S., Noakes, D. L. G., Thompson, N., Blouin, M., Morrison, B., Schreck, C. B. 2013.  Exposure to elevated temperature during early development affects sexual development in Oncorhynchus mykiss. North Pacific Anadromous Fish Commission Technical Report No. 9: 101-103.

Cole, K.S., Noakes, D.L.G., Thompson, N., Blouin, M., Morrison, B., Couture, R.B., O’Neil, J. and C.B. Schreck. 2021. Effects of temperature on sexual development in steelhead, Oncorhynchus mykiss. Environmental Biology of Fishes, pp. 1-10.

Cooke, S.J., Auld, H.L., Birnie-Gauvin, K., Elvidge, C.K., Piczak, M.L., Twardek, W.M., Raby, G.D., Brownscombe, J.W., Midwood, J.D., Lennox, R.J., and C. Madliger. 2022. On the relevance of animal behavior to the management and conservation of fishes and fisheries. Environmental Biology of Fishes, pp.1-26.

Couch, C.E., Colvin, M., Chitwood, R., Peterson, J.T., and C.B. Schreck. 2022. Scope of the cortisol stress response in Chinook salmon during maturation. Fisheries Research. 254:106416.

Couch, C.E., Kent, M., Weiss, L., Cummings, L., Takvorian, P., and Sanders, J. 2022. Enterocytozoon schreckii, n.sp., infects the enterocytes of adult Chinook salmon and may be a sentinel of immunosenescence. mSphere. 7(1): e00908-21.

Couch, C.E., Neal, W.T., Herron, C.L., Kent, M.L., Schreck, C. B., and J.T. Peterson. 2023. Gut microbiome composition associates with corticosteroid treatment, morbidity, and senescence in Chinook salmon (Oncorhynchus tshawytscha). Scientific Reports, 13(1), 2567.

Dayan, D.I., Lorion, C., and K.G. O’Malley. 2023. Population structure of Oregon coastal Chum Salmon (Oncorhynchus keta) and considerations for carcass tissue sampling. Science Bulletin 2023‐07. Oregon Department of Fish and Wildlife, Salem. 28 pp.

Dayan, D.I., Mazur, S., Green, L.J., Bohn, S., Johnson, M.A., Van Dyke, D.J., Samarin, P.A., Battleson, R.D., and K.G. O’Malley. 2023. Genetic diversity within late-summer and half-pounder steelhead (Oncorhynchus mykiss) in the Rogue River, Oregon. Conservation Genetics.

Dayan, D.I., Sard, N.M., Johnson, M.A., Fitzpatrick, C.K., Couture, R., and K.G. O’Malley. 2024. A singe generation in the wild increases fitness for descendants of hatchery-origin Chinook salmon (Oncorhynchus tshawytscha). Evolutionary Applications 2024;17:e13678

Dittman, A.H., Cunningham, C.J., and T.P. Quinn. 2022.Can unique amino acid profiles guide adult salmon to natal streams? A comparison of streams sampled prior to and after the arrival of adult Pacific salmon. Hydrobiologia 849:3501–3513.

Dittman, A.H., May, D., Johnson, M., Baldwin, D., Scholz, N., and J. Athos. 2024. Odor exposure during imprinting periods increases odorant-specific sensitivity and receptor gene expression in coho salmon (Oncorhynchus kisutch). Journal of Experimental Biology jeb.247786.

Dittman, A.H., Pearsons, T.N., May, D., Couture, R.B., and D.L.G Noakes. 2015. Imprinting of hatchery-reared salmon to targeted spawning locations: a new embryonic imprinting paradigm for hatchery programs. Fisheries, 40(3), 114-123.

Evans, M.L., Hard, J.J., Black, A.N., Sard, N.M., and K.G. O’Malley. 2019. A quantitative genetic analysis of life-history traits and lifetime reproductive success in reintroduced Chinook salmon. Conservation Genetics 20, 781-799.

Evans, M.L., Johnson, M.A., Jacobson, D., Wang, J., Hogansen, M. and O’Malley, K.G., 2016. Evaluating a multi-generational reintroduction program for threatened salmon using genetic parentage analysis. Canadian Journal of Fisheries and Aquatic Sciences, 73(5), pp.844-852.

Ford, M., Nichols, K., Waples, R., Anderson, E., Kardos, M., Koch, I., McKinney, G., Miller, M., Myers, J., Naish, K,, Narum, S., O’Malley, K.G., Pearse, D., Seamons, T., Spidle, A., Swanson, P., Thompson, T., Warheit, K., and S. Willis. 2020. Reviewing and synthesizing the state of the science regarding associations between adult run timing and specific genotypes in Chinook salmon and steelhead. Report of a workshop held in Seattle, WA. 27-28 February 2020. 42 pp

Gao, Y. and Noakes, D.L., 2012. Chemical signatures of otoliths and application in fisheries. Environmental Biology of Fishes, 95(4), pp.415-418.

Herron C.L., Cogliati K.M., Dolan B., Munakata A., Schreck CB. 2018. Stress up-regulates oxidative burst in juvenile Chinook salmon leukocytes. Fish & Shellfish Immunology 80:655-659.

Griffiths R., Noakes D.L.G. 2021. Effect of eroded sediments from land-slip on the benthic macroinvertebrate community in Fall Creek. Report to the OHRC Board. 10 pp.

Hahlbeck, N.H., Anlauf-Dunn, K.J., Piotrowski, S.J., Ortega, J.D., Tinniswood, W.R., Eliason, E.J., O’Malley, K.G., Sloat, M.R., Wyatt, M.A., Hereford, M., Ramirez, B.S., and J.B. Armstrong. 2023. Habitat fragmentation drives divergent survival strategies of a cold-water fish in a warm landscape. Ecosphere, 14(7), e4622.

Havey, M.A., Dittman, A.H., Quinn, T.P., Lema, S.C., and D. May. 2017. Experimental evidence for olfactory imprinting by Sockeye salmon at embryonic and smolt stages. Transactions of the American Fisheries Society. 146:74-83.

Herron, C.L., Cogliati, K.M., Dolan, B., Munakata, A., and C.B. Schreck. 2018. Stress up-regulates oxidative burst in juvenile Chinook salmon leukocytes. Fish & Shellfish Immunology 80:655-659.

Herron, C.L., Cogliati, K.M., Krajcik, J.A., Noakes, D.L.G., Peterson, J.T., and C.B. Schreck. 2023. Effects of rearing density and in-tank structure on the stress response in juvenile Chinook salmon (Oncorhynchus tshawytscha). Aquaculture Reports, 33, 101876.

Howell, M. 2021. A performance evaluation of in-stream hatchboxes and streamside incubators for Chinook salmon. BSc thesis for BioResource Research at OSU. 20 pp.

Jensen, A., Schreck, C.B., Hess, J., North, J., Bohn, S., O’Malley, K.G., and J.T. Peterson. 2020. Application of genetic stock identification and parentage‐based tagging in a mixed‐stock recreational Chinook salmon fishery. North American Journal of Fisheries Management 41(1): 130-141.

Johnson, M.A. and Friesen, T.A., 2013. Age at maturity, fork length, and sex ratio of upper Willamette River hatchery spring Chinook salmon. North American Journal of Fisheries Management, 33(2), pp.318-328.

Johnson, M.A., Friesen, T.A., Van Doornik, D.M., Teel, D.J. and J.M. Myers. 2021. Genetic interactions among native and introduced stocks of Oncorhynchus mykiss in the upper Willamette River, Oregon. Conservation Genetics 22:111-124.

Johnson, M.A., Friesen, T.A., VanDoornik, D.M., Teel, D.J., and J.M. Myers. 2018. Genetic influence from hatchery stocks on upper Willamette River steelhead Oncorhynchus mykiss. ODFW Information Report 2018-03, 20 pp.

Johnson, M.A., Jones, M.K., Falcy, M.R., Spangler, J., Couture, R.B. and D.L.G. Noakes. 2023. Can angler-assisted broodstock collection programs improve harvest rates of hatchery-produced steelhead? Environmental Biology of Fishes 106:1079–1092.

Johnson, M.A., Noakes, D.L.G., Friesen, T.A., Dittman, A.H., Couture, R.B., Schreck, C.B., Banner, C., May, D., and T.P. Quinn. 2019. Growth, survivorship, and juvenile physiology of triploid steelhead (Oncorhynchus mykiss). Fisheries Research 220.

Johnson, M.A., Spangler, J., Jones, M., Couture, R.B., and D.L.G Noakes. 2020. Angler harvest of Alsea River hatchery winter steelhead: An evaluation of wild broodstock collection techniques. ODFW Information Report no. 2020-05. 43 pp.

Klimley, A.P., Putman, N.F., Keller, B.A., and D.L.G. Noakes. 2021. A call to assess the impacts of electromagnetic fields from subsea cables on the movement ecology of marine migrants. Conservation Science and Practice, 3(7), e436.

Koeberle, A.L., Arismendi, I., Crittenden, W., Leer, D. and D.L.G. Noakes. 2020. Fluctuating asymmetry of adult Chinook salmon (Oncorhynchus tshawytscha) otoliths from wild and hatchery origins. Aquatic Ecology 54:431-446.

Leblanc, C.A., Schreck, C., Kristjánsson, B.K., Skúlason, S. and D.L.G. Noakes. 2022. Egg size–related traits during the first year of growth and smolting in hatchery and wild juveniles of steelhead trout Oncorhynchus mykiss. Environmental Biology of Fishes, pp.1-18.

Mani, A., Henn, C., Couch, C., Patel, S., Korytar, T., and I. Salinas. 2023. A niche-adapted brain microbiome in salmonids at homeostasis. bioRxiv, 2023-12.

Minkoff, D., Ardren, W., Kaiser, K., Dittman, A.H., Quinn, T., Atema, J., and B. Taylor. 2023. Dissolved free amino acids could be odorants for imprinting and homing by Atlantic Salmon. Freshwater Science 42(4):375-391.

Munakata, A., Ogihara, E., Schreck, C.B., and D.L.G. Noakes. 2017. Effects of short term acclimation in cool and warm water and influent water temperatures on temperature selection behavior in juvenile steelhead trout, Oncorhynchus mykiss. Aquaculture 467: 219-224.

Naisbett-Jones, L.C., Putman, N.F., Scanlan, M.M., Noakes, D.L.G., and K.J. Lohmann. 2020. Magnetoreception in fishes: the effect of magnetic pulses on orientation of juvenile Pacific salmon. Journal of Experimental Biology, 223(10).

Obley, M. 2023. Modifications to rearing environments as a method for reducing domestication selection in hatchery-reared steelhead (Oncorhynchus mykiss). MSc thesis in Integrative Biology. Oregon State University.

O’Malley, K.G., Bohn, S., and C.K. Fitzpatrick. 2021. Adult salmonids trap and transport success above dams. Technical report submitted to the U.S. Army Corps of Engineers.

O’Malley, K.G., Bohn, S., Whitman, L., and M. Lewis. 2020. Genetic analysis of Chinook salmon in the Clackamas Basin, 2015-2018. Information Report 2020-04. Oregon Department of Fish and Wildlife, Salem. 20pp.

O’Malley, K.G., Dayan, D.I., Fitzpatrick, C.K., and R. Couture. 2023. Evaluating spring Chinook salmon releases above Cougar Dam, on the South Fork McKenzie River, using genetic parentage analysis. Technical report submitted to the U.S. Army Corps of Engineers. Portland, OR.

O’Malley, K.G., Dayan, D.I., Fitzpatrick, CK, Bohn, S., and G. Grenbemer. 2023. Evaluating spring Chinook salmon releases above Detroit Dam and below Big Cliff Dam, on the North Santiam River, using genetic parentage analysis. Technical report submitted to the U.S. Army Corps of Engineers. Portland, OR.

O’Malley, K.G., Dayan, D.I., Mazur, S., Green, L.J., Fitzpatrick, C.K., Bohn, S., and A. Wells. 2022. Evaluating the genetics of naturally produced Chinook salmon (Oncorhynchus tshawytscha) captured in the Lower Rogue River (OR) in 2020. Science Bulletin 2022-09. Oregon Department of Fish and Wildlife, Salem.

O’Malley, K.G., Hereford, M.E., Olsen, K.C., Pearse, D.E., Tinniswood, W.R., Ramirez, B.S., Armstrong, J.B., and S. Piotrowski. In Prep. Characterizing fine-scale neutral and adaptive genetic diversity of Oncorhynchus mykiss in the Klamath Basin before dam removal.

O’Malley, K.G., Fitzpatrick, C.K., Olsen, K.C., and R. Couture. 2024. Evaluating spring Chinook releases above Foster Dam, on the South Santiam River, using genetic parentage analysis. Technical report submitted to the U.S. Army Corps of Engineers. Portland, OR.

O’Malley, K.G., Fitzpatrick, C.K., Bohn, S., Van Dyke, D., Samarin, P., Pease, D., and M. Semrow. 2022. Identification of homozygous fall run Chinook salmon (Oncorhynchus tshawytscha) returning to Cole Rivers Hatchery on the Rogue River (OR). Science Bulletin 2022-08. Oregon Department of Fish and Wildlife, Salem.

O’Malley, K.G., Mazur, S., Green, L.J., Bohn, S., and A. Wells. 2020b. Evaluating the genetics of naturally produced Chinook salmon in the lower Rogue River fishery. Information Report 2020-02. Oregon Department of Fish and Wildlife, Salem. 11 pp.

O Malley, K.G., Van Dyke, D., Samarin, P.A., and S. Bohn. 2019. Evaluating the genetics of Rogue River Chinook salmon. Technical Report to the U.S. Army Corps of Engineers.

O’Malley, K.G., Van Dyke, D., Samarin, P.A., Bohn, S., and S. Clements. 2020c. An evaluation of “early” and “late” run alleles in Rogue River Chinook salmon (Oncorhynchus tshawytscha). Information Report 2020-06. Oregon Department of Fish and Wildlife, Salem. 24 pp.

Pollock, A.M., Kamran, M., Dittman, A.H., Johnson, M.A., and D.L.G. Noakes. 2020. Within-river straying: sex and size influence recovery location of hatchery Chinook salmon (Oncorhynchus tshawytscha). Canadian Journal of Fisheries and Aquatic Sciences 77(2), 226-235.

Pope, A.C., Kock, T.J., Perry, R.W., Cogliati, K.M., O’Malley, K.G., Murphy, C.A., and S.D. Fielding. In Revision. Using parentage based tagging to estimate survival of Chinook salmon fry in a large reservoir. Environmental Biology of Fishes 107, 735–754.

Putman, N.F., Scanlan, M.M., Pollock, A.M., O’Neil, J.P., Couture, R.B., Stoner, J.S., Quinn, T.P., Lohmann, K.J., and D.L.G. Noakes. 2018. Geomagnetic field influences upward movement of young Chinook salmon emerging from nests. Biology Letters 14(2): 20170752.

Putman, N. F., Ueda, H., and D.L.G. Noakes. 2019. The current status of research on geomagnetic navigation in Pacific salmon. North Pacific Anadromous Fish Commission Technical Report 15: 1–5.

Putman, N.F., Williams, C.R., Gallagher, E.P., and A.H. Dittman. 2020. A sense of place: pink salmon use a magnetic map for orientation. Journal of Experimental Biology, 223(4), jeb218735.

Putman, N.F., Scanlan, M.M., Billman, E.J., O’Neil, J.P., Couture, R.B., Quinn, T.P., Lohmann, K.J. and Noakes, D.L., 2014. An inherited magnetic map guides ocean navigation in juvenile Pacific salmon. Current Biology, 24(4), pp.446-450.

Rand, P.S., Berejikian, B.A., Pearsons, T.N. and Noakes, D.L., 2012. Ecological interactions between wild and hatchery salmonids: an introduction to the special issue. Environmental Biology of Fishes, 94(1), pp.1-6.

Reed, A.N., Rowland, F.E., Krajcik, J.A., and D.E. Tillitt. 2024. Thiamine Supplementation Improves Survival and Body Condition of Hatchery-Reared Steelhead (Oncorhynchus mykiss) in Oregon. Veterinary Science 10(2):156.

Sard, N. M., M. A. Johnson, D. P. Jacobson, M. J. Hogansen, K. G. O’Malley and M. A. Banks. 2016. Genetic monitoring guides adaptive management of a migratory fish reintroduction program. Animal Conservation 19(6): 570-577.

Sard, N. M., K. G. O’Malley, D. P. Jacobson, M. J. Hogansen, M. A. Johnson and M. A. Banks 2015. Factors influencing spawner success in a spring Chinook salmon (Oncorhynchus tshawytscha) reintroduction program. Canadian Journal of Fisheries and Aquatic Sciences 72:1390-1397

Scanlan, M., Putman, N.F., Pollock, A.M., and D.L.G. Noakes. 2018. Magnetic map in nonanadromous Atlantic salmon. Proceedings of the National Academy of Science.

Self, K.E., Schreck, C.B., Cogliati, K.M., Billman, E.J., and D.L.G. Noakes. 2018. Egg size and growth in steelhead Oncorhynchus mykiss. Journal of Fish Biology 93(3): 465 – 468.

Self, K.E., Schreck, C.B., Cogliati, K.M., Billman, E.J., and D.L.G. Noakes. 2018. The effect of rearing structures on behaviour and movement of juvenile steelhead Oncorhynchus mykiss. Journal of Fish Biology 93(3): 449 – 454.

Stewart, H.A., Cogliati, K.M., Billman, E.J., Chitwood, R., Unrein, J.R., Noakes, D.L.G., and C.B. Schreck. 2017. Effects of transportation timing on osmoregulation and survival in yearling hatchery Chinook salmon (Oncorhynchus tshawytscha). Journal of Applied Aquaculture 29(3-4): 277-290.

Thompson, N.F. and M.S. Blouin.2016. Family dominance level measured during the fry stage weakly influences family length at smolting in hatchery reared steelhead (Oncorhynchus mykiss). Transactions of the American Fisheries Society 145: 1282-1289.

Thompson, N.F. and M.S. Blouin. 2015. The effects of high rearing density on the potential for domestication selection in hatchery culture of steelhead (Oncorhynchus mykiss). Canadian Journal of Fisheries and Aquatic Sciences 72:1-6.

Thompson, N.F., Clemens, B., Ketchum, L., Simpson, P., Reagan, R. and M.S. Blouin. 2018. Family influence on length at release and size-biased survival post release in hatchery-reared steelhead: a mechanism to explain how domestication occurs. Aquaculture 491:135-146.

Unrein, J.R., Billman, E.J., Cogliati, K.M., Chitwood, R. Noakes, D.L.G., and C.B. Schreck. 2018. Vertical self-sorting behavior in juvenile Chinook salmon (Oncorhynchus tshawytscha): evidence for family differences and variation in growth and morphology. Environmental Biology of Fishes, 101(2): 341-353.

Vaux, F., Bohn, S., Hyde, J.R., and K.G. O’Malley. 2021. Adaptive markers distinguish North and South Pacific Albacore amid low population differentiation. Evolutionary Applications.

Waples, R.W., Ford, M.J., Nichols, K., Kardos, M., Myers, J., Thompson, T.Q., Anderson, E.C., Koche, I.J., McKinney, G., Miller, M.R., Naish, K., Narum, S.R., O’Malley, K.G., Pearse, D., Pess, G.R., Quinn, T.P., Seamons, T.R., Spidle, A., Warheit, K., and S.C. Willis. 2022. Implications of large-effect loci for conservation: a review and case study with Pacific salmon. Journal of Heredity 113(2):121-144.

White, S.M., Dittman, A.H., Johnson, M.A., and T.P. Quinn. 2024. Climate-driven straying dynamics in anadromous salmon and steelhead: Research agenda for conservation. Ecology of Freshwater Fish 2024;00:e12797.

Whitmore, M., Richardson, S., Huff, A., Goodson, K., Quinn, T.P., Dittman, A.H., Johnson, M.A., Kamran, M., and D.L.G Noakes. 2021. Homeward bound: In‐river movements of adult hatchery‐and natural‐origin Chinook Salmon in the Elk River, Oregon. North American Journal of Fisheries Management 41(4): 1088-1096.

Wrey, M.C., Skaar, J.C., Cates, S.B., Bollmann, S.R., Fuentes, C., and M.S. Blouin. 2020. Family variation in surface feeding behavior of steelhead fry predicts growth rate under hatchery conditions. bioRxiv doi: 10.1101/2020.08.07.241141